
This marine lab is located about one hundred feet from the shore of Harpswell Sound. Large windows along the ocean side afford views of the Sound and gorgeous sunsets, both seen through the intervening forest.
The lab is equipped with running sea water and currently there are sixteen aquaria available for research and teaching. There is one large flow tank for studying the effects of flow on marine organisms and plans call for two additional flow tanks and a large wave tank. In addition, four special large tables with sea water aquaria insets aid in the observation of marine life. There is a computer work space and plans call for a discussion/seminar room. Biology, earth and oceaographic science and environmental studies courses use the lab. In addition students and faculty have ongoing research projects in this lab.

Summer 2012
Holly Blackburn, William and Mary College
Advisor: Jonathan Allen (William and Mary College)
Project: The Effect of Maternal Investment on Cloning Frequency in the Sea Star Asterias forbesi
Anna Chase, '13
Advisor: Trevor Rivers, Award: Doherty Fellowship
Project: Effects of Photopollution on hte Settlement of Marine Fouling Communities in the Gulf of Maine
Katherine Guttenplan '13
Advisor: Amy Johnson, Award: Howard Hughes Medical Institute Postbac Summer Fellowship
Project: The Effects of climate change on the growth of the sea urchin Strongylocentrotus droebachiensis
Julia Livermore '13
Advisor: Trevor Rivers, Award: Doherty Fellowship
Project: Effects of photopollution on the settlement of marine fouling communitees in the Gulf of Maine
Tamara Perreault '12
Advisor: Trevor Rivers, Award: Howard Hughes Medical Institute Postbac Summer Fellowship
Project: Predation Response Behavior of the Luminescent Scale Worm Harmothoe imbricata
Mark Smithson, William and Mary College
Advisor: Jonathan Allen
Project: Size-based differences in performance of Strongylocentrotus drobachiensis
Shelby Ziegler, College of William and Mary
Advisor: Jonathan Allen
Project: Rates and costs of cloning in Echinoderms
Summer 2011
Holly Blackburn, College of William and Mary
Advisor: Jonathan Allen
Project: Tricking the embryo: Blastomere separations in deuterostomes
Anna Chase, ‘13
Advisor: Amy Johnson, Award: Doherty Fellowship
Project: Effects of Temperature on Growth Rate of the Green Sea Urchin, Strongylocentrotus droebachiensis
Katherine Guttenplan, '12
Advisor: Amy Johnson, Award: Doherty Fellowship
Project: The Effects of Climate Change on the Growth of the Sea Urchin Strongylocentrotus Droebachiensis
Kelly Hoolihan, College of William and Mary
Advisor: Jonathan Allen
Project: Developmental plasticity and invasive species
Gina Lonati '12
Advisor: Amy Johnson, Award: Doherty Fellowship
Project: Using the green sea urchin, Stronglyocentrotus, as an environmental indicator on its response to global climate change
Tamara Perreault '12
Advisor: Trevor Rivers, Award: Doherty Fellowship
Project: Luminescence as a response to predators in the scale worm Harmothoe imbricate
Jordan Salyers, College of William and Mary
Advisor: Jonathan Allen
Project: Estimating Predation Rates in Juvenile Sea Urchin
Daniel Schwab '12, College of William and Mary
Advisor: Jonathan Allen
Project: The effect of maternal size on reproduction and larval development in the mud snail Ilyanassa
Summer 2010
John Roger Brothers, '11
Advisor: Amy Johnson and Dan Thornhill, Award: HHMI
Effects of Climate Change on the Growth and Calcification of the Green Sea Urchin, Strongylocentrotus droebachiensis
Connor Gallagher, '12
Advisor: Jonathan Allen and Amy Johnson, Award: HHMI
Ocean Acidifications Effect in Marine Calicifier Skeletal Structure and Predation Rates
Laura Newcomb, '11
Advisor: Daniel Thornhill and Amy Johnson, Award:Doherty Fellowship
The Effects of Climate Change on the Calfification of the Temperate Coral Astrangia poculata
Jordan Salyers, College of William and Mary
Advisor: Jonathan Allen
Estimating Predation Rates in Juvenile Sea Urchin
Connor White, College of William and Mary
Advisor: Jonathan Allen
The effect of egg capsule clustering on embryonic survival in the dogwhelk Nucella lapillus
Summer 2009
Alexandria Brasili ‘10
Advisor: Amy Johnson, Award: Rusack Fellowship
Temperature and growth in the green sea urchin Strongylocentrotus droebachiensis
Jill Dixon '09, Randolph Macon College
Advisor: Jonathan Allen
The role of encapsulation in the marine gastropod, Nucella lapillus
Katharine Doubleday ‘11
Advisor: Dan Thornhill, Award: Doherty Fellowship
Effects of coral hybridization on specificity of coral-Symbiodinium symbiosis in Caribbean reef-building corals
Laura Newcomb '11
Advisor: Dan Thornhill, Award: Doherty Fellowship
Investigating the Nature of a Temperate Coral Symbiosis Coral
Emily Norton '10
Advisor: Amy Johnson, Award: Doherty Fellowship
Mathematical modeling of underwater walking in three species of intertidal crabs: Carcinus maenas, Hemigrapsus sanguineus, and Cancer irroratus
Amanda Santoni, Randolph Macon College
Advisor: Jonathan Allen
Predator induced plasticity in maternal investment of the mud snail Ilyanassa obsoleta
TENTH SUMMER
Summer 2008/Academic Year 2008-2009 (Johnson sabbatical thru summer 2008)
(1) Rachel Dicker (’09) (Doherty Fellow, Summer 2008; Independent research, Academic year 2008-2009). Rachel studied egg capsule distribution in Nucella lapillus as a strategic maternal investment and the distribution of stacks of the slipper shell Cripedula fornicata with direction from 2005-2008 Doherty Marine Biology Scholar Jonathan Allen (Summer 2008) and Professor Amy Johnson (Academic year 2008-2009)
(2) Ashton Bunce (Juniata College in Huntington, PA, Summer 2008). Ashton worked with Rachel Dicker and 2005-2008 Doherty Marine Biology Scholar Jonathan Allen contributing to research in maternal investment in egg capsule distribution in Nucella lapillus.
NINTH SUMMER
Summer 2007/Academic Year 2007-2008 (Johnson sabbatical thru summer 2008)
(1) Nick Alcorn (’08). (Doherty Fellow, Summers 2006, 2007; Senior Honors research, Academic year 2007-2008). Nick worked with 2005-2008 Doherty Marine Biology Scholar Jonathan Allen on (1) Egg size, parental investment, and larval development in the sand dollar Echinarachnius parma, (2) the effects of maternal investment on the development and growth of larval green sea urchins, and (3) fluorochrome labeling of juvenile urchins. Nick won a best paper award for a poster of his work at the annual meeting of the Society of Integrative and Comparative Biology in January, 2008. Senior Honors Research Title: How do changes in parental investment influence larval development in Gulf of Maine echinoids?
(2) Ian Haight (’08). (Howard Hughes Fellow, Summers 2006, 2007, Academic year 2007-2008). Ian examined the effects of temperature changes on the larval growth and development of two local seastar species (Asterias rubens and Asterias forbesi) as well as the hybrids that frequently form between them under the guidance of 2005-2008 Doherty Marine Biology Scholar Jonathan Allen and 2003-2004 Coastal Studies Center Scholar-in-Residence and visiting Assistant Professor Marney Pratt (’97).
(3) Julia Kleederman (Wesleyan University, Summer 2007). Julia worked on several projects with 2005-2008 Doherty Marine Biology Scholar Jon Allen, including measurements of hermit crab feeding preferences.
(4) Kat Anderson (’08). (Senior Honors research, Academic year 2007-2008). Senior honors project title: Comparing chemical and structural induced defenses of Ascophyllum nodosum and Fucus vesiculosus in response to damage by herbivory and wave action with visiting assistant professor Kurt Bretsch.
EIGTH SUMMER
Summer 2006/Academic Year 2006-2007
(1) Nick Alcorn (’08). (Doherty Fellow, Summers 2006, 2007; Senior Honors research, Academic year 2007-2008). Nick worked with 2005-2008 Doherty Marine Biology Scholar Jonathan Allen on (1) Egg size, parental investment, and larval development in the sand dollar Echinarachnius parma, (2) the effects of maternal investment on the development and growth of larval green sea urchins, and (3) fluorochrome labeling of juvenile urchins. Nick won a best paper award for a poster of his work at the annual meeting of the Society of Integrative and Comparative Biology in January, 2008. Senior Honors Research Title: How do changes in parental investment influence larval development in Gulf of Maine echinoids?
(2) Ian Haight (’08). (Howard Hughes Fellow, Summers 2006, 2007, Academic year 2007-2008). Ian examined the effects of temperature changes on the larval growth and development of two local seastar species (Asterias rubens and Asterias forbesi) as well as the hybrids that frequently form between them under the guidance of 2005-2008 Doherty Marine Biology Scholar Jonathan Allen and 2003-2004 Coastal Studies Center Scholar-in-Residence and visiting Assistant Professor Marney Pratt (’97).
(3) Rebecca Selden (’06). (Beckman Fellow and Senior Honors Research, Summer 2005 through Summer 2006). Now in graduate school at U.C. Santa Barbara. Becca was a recipient of the prestigious Beckman fellowship for her work on the influence of predator scent on growth sea urchins under the guidance of Professor Amy Johnson and with the assistance of Research Associate Olaf Ellers. This research is now published: http://www.springerlink.com/content/100441/?k=selden. Senior Honors Research Title:: Growth and Metabolism of the Green Sea Urchin, Strongylocentrotus droebachiensis: Flow environments and predation pressure.
(4) Eric Robinson (’07). (Howard Hughes Fellow, Summer 2006). Eric continued his summer research in the marine lab with a two semester independent study in the marine lab measuring the palatability of marine invertebrate embryos to predators including lobsters, green crabs, and rock crabs under the guidance of 2005-2008 Doherty Marine Biology Scholar Jon Allen
(5) Bobbi Robinson (’08). (Doherty Fellow, Summer 2006). Bobbi worked on feeding strategies and growth patterns of two species of sea stars (Asterias rubens and Asterias forbesi) in relation to various parameters including temperature, competition and cannibalism under the guidance of 2003-2004 Coastal Studies Center Scholar-in-Residence and visiting Assistant Professor Marney Pratt (’97).
(6) Orissa Moulton (Wheaton College, Summer 2006), worked with 2003-2004 Coastal Studies Center Scholar-in-Residence and visiting Assistant Professor Marney Pratt(’97), Bobbi Robinson (’08) and Ian Haight (’08) on the feeding strategies and growth patterns of two species of sea stars (Asterias rubens and Asterias forbesi) in relation to various parameters including temperature, competition and cannibalism.
(7) Ben Lake (’07). (Kent Island Summer Research Fellow, Hughes Award, Summer, 2006, and INBRE award for Senior Honors research, Bowdoin Marine Lab, Academic year 2006-2007). Ben measured metabolic and feeding rates of barnacles in different orientations to flow under the guidance of 2004-2008 Bowdoin Scientific Station Director and Associate Professor of Biology at Kenyon College Bob Mauck and Bowdoin Professor Amy Johnson. Ben received an award for best student poster for his presentation of his research at the annual Society of Integrative and Comparative Biology meeting in January, 2007 in Phoenix, Arizona. Senior Honors Research Title: The influence of orientation and water velocity on the feeding behavior and metabolism of Semibalanus balanoides.
SEVENTH SUMMER
Summer 2005/Academic Year 2005-2006
(1) Rebecca Selden (’06). (Beckman Fellow and Senior Honors Research, Summer 2005 through Summer 2006). Now in graduate school at U.C. Santa Barbara. Becca was a recipient of the prestigious Beckman fellowship for her work on the influence of predator scent on growth sea urchins under the guidance of Professor Amy Johnson and with the assistance of Research Associate Olaf Ellers. This research is now published: http://www.springerlink.com/content/100441/?k=selden. Senior Honors Research Title:: Growth and Metabolism of the Green Sea Urchin, Strongylocentrotus droebachiensis: Flow environments and predation pressure.
(2) Molly Wright (‘05). (Rusack Fellow, Summer 2004; INBRE Postgraduate Fellow, Summer 2005, Senior Honors Research, Academic year 2004-2005). Now in graduate school at U.C. Berkeley, Molly studied growth and metabolism in sea urchins under the guidance of Professor Amy Johnson and Research Associate in Mathematics and Biology Olaf Ellers. Molly presented her senior honors research at the annual meeting of the Society of Integrative and Comparative Biology in Toronto in 2004. Senior Honors Research Title: Growth and metabolism in the green sea urchin, Strongylocentrotus droebachiensis.
(3) Kiersa Benson (‘06). (Doherty Fellow, Summer 2005; Senior Honors research, Academic year 2006-2007). Kiersa conducted field and laboratory experiments in the summer of 2006 under the guidance of visiting Assistant Professor Lindsay Whitlow, followed by a five-month growth study conducted in the marine lab, on a hard-shell clam under the guidance of Professor Amy Johnson. Senior Honors Research Title: Growth and behavioral response of hard shell clam Mercenaria mercenaria in response to crab predation cues
SIXTH SUMMER
Summer 2004/Academic Year 2004-2005
(1) Peter Benjamin Stranges (‘05). (Doherty Fellow, Summer 2004; Senior Honors Research, Academic year 2004-2005). Ben explored the mathematical possibilities of modeling von Bertalanffy growth in sea urchins with a variable exponent that relied on oxygen transport through boundary layers under the guidance of Professor Amy Johnson and Research Associate in Mathematics and Biology Olaf Ellers. Now in graduate school in biophysics at the University of North Carolina, Ben presented his senior honors research at the annual meeting of the Society of Integrative and Comparative Biology in Toronto in 2004. Senior Honors Research Title: Boundary layer fluid dynamics effects on oxygen metabolism and growth rates in green sea urchins, Strongylocentrotus droebachiensis.
(2) Molly Wright (‘05). (Rusack Fellow, Summer 2004; INBRE Postgraduate Fellow, Summer 2005, Senior Honors Research, Academic year 2004-2005). Now in graduate school at U.C. Berkeley, Molly studied growth and metabolism in sea urchins under the guidance of Professor Amy Johnson and Research Associate in Mathematics and Biology Olaf Ellers. Molly presented her senior honors research at the annual meeting of the Society of Integrative and Comparative Biology in Toronto in 2004. Senior Honors Research Title: Growth and metabolism in the green sea urchin, Strongylocentrotus droebachiensis.
FIFTH SUMMER
Summer 2003/Academic Year 2003-2004
Eucidaris tribuloides spines with respect to muscle and ligament properties.
(1) Lynne Davies (’04). (Doherty Fellow, Summer 2003; Senior Honors research, Academic year 2003-2004). Lynne studied the relationship between growth and echinoderm catch connective tissue under the guidance of Professor Amy Johnson and Research Associate in Mathematics and Biology Olaf Ellers. Senior Honors Research Title: An analysis of diet, spine length, competition and mutable collagenous tissues as indicators of somatic and gonadal growth in the green sea urchin, Strongylocentrotus droebachiensis.
(2) Benjamin Ming Fei Chan (‘05). (Gibbons Fellow, Summer 2003; Senior Honors research, Academic year 2004-2005). Ben studied echinoderm catch connective tissue under the guidance of Professor Amy Johnson and Research Associate in Mathematics and Biology Olaf Ellers. Now in dental school. Senior Honors Research Title: Exploring tensile attributes of mutable collagenous tissue in
(3) Angela Senese (’04). (Senior Honors research, Academic year 2003-2004). Angie used the flow tanks in the marine lab to study the hydrodynamics of schooling in fish under the guidance of Professor Amy Johnson. Senior Honors Research Title: The hydrodynamics of schooling behavior in the Marsh Killifish, Fundulus heteroclitis.
(4) Emily Grason (‘04). (Senior Honors research, Academic year 2003-2004). Emily studied aspects of the ecology of an invasive species of bryozoan under the guidance of 2003-2004 Coastal Studies Center Scholar-in-Residence and visiting Assistant Professor Marney Pratt (’97). Her work has been published: http://www.springerlink.com/content/a41748360j835110/ Senior Honors Research Title: Influence of an invasive bryozoan on Onchidoris muricata prey selection in the Gulf of Maine.
(5) Jon Harris (‘05). (Hughes Fellow, Summer 2003). Jon examined the effects of Littorina littorea (periwinkles) on algal and green crab populations in the rocky intertidal in and around the Coastal Studies Center under the guidance of visiting Assistant Professor Lindsay Whitlow.
FOURTH SUMMER
Summer 2002/Academic Year 2002-2003
(1) Laura Windecker (‘03). (Doherty Fellow, Summer 2002). Laura studied diet and growth in sea urchins, tested mathematical models of growth and studied catch connective tissue under the guidance of Professor Amy Johnson and Research Associate in Mathematics and Biology Olaf Ellers. Laura presented the results of her Senior Honors research at the annual meeting of the Society of Integrative and Comparative Biology in 2002. Laura is in a graduate program in Oceanography at the University of Rhode Island. Senior Honors Research Title: Mechanisms of size-specific growth in the sea urchin, Strongylocentrotus droebachiensis.
(2) Kara Podkaminer (’03). (Senior Honors research, Academic year 2002-2003). Kara used the flow tanks in the marine lab to study the hydrodynamics of schooling in fish under the guidance of Professor Amy Johnson. Senior Honors Research Title: Hydrodynamic benefits of schooling as measured by tail beat and opercular frequency in killifish, Fundulus heteroclitus.
(3) Dave Donnelly (‘03). (Hughes Fellow, Summer 2002). Dave studied zooplankton grazing, harmful algal blooms and toxic accumulation kinetics in planktonic food webs under the guidance of visiting Assistant Professor Greg Teegarden.
(4) Stephen Carpenter (‘03). (Rusack Fellow, Summer 2002). Steve studied zooplankton grazing, harmful algal blooms and toxic accumulation kinetics in planktonic food webs with visiting Assistant Professor Greg Teegarden.
(5) Louis Plough (’03). (Senior Honors research, Academic year 2002-2003). Louis studied the effect of predators on the behavior of burrowing clams under the guidance of visiting Assistant Professor Lindsay Whitlow. Senior Honors Research Title: The effects of green crab Carcinus maenas predatory cues on the burrowing depth and morphology of the soft-shelled clam Mya arenaria.
(6) Elizabeth Bryson (’03). (Senior Honors research, Academic year 2002-2003). Elizabeth studied the effect of predators on the behavior of burrowing clams under the guidance of visiting Assistant Professor Lindsay Whitlow. She went on to graduate school at Northeastern University in Boston. Senior Honors Research Title: The effects of crab cues and clam density on Mya arenaria : burrowing depth and resource allocation.
(7) Michelle Weaver (‘03). (Gibbons Fellow, Summer 2001, 2002). Michelle worked on the Coastal Studies Center Marine Organism database under the guidance of Laboratory Instructory David Guay.
THIRD SUMMER
Summer 2001/Academic Year 2001-2002 (Johnson sabbatical thru summer 2001)
(1) Dean Anson (‘03). (Rusack Fellow, Summer 2001). Dean worked on toxic algal blooms with visiting Assistant Professor Greg Teegarden. He is a coauthor on a publication of this work (Copepod feeding response to varying Alexandrium spp. cellular toxicity and cell concentration among natural plankton samples; doi:10.1016/j.hal.2007.05.010)
(2) Stacy Barron (‘03). (Hughes Fellow, Summer 2001). Stacy worked on Phycotoxin accumulation in zooplankton feeding on Alexandrium fundyense under the guidance of visiting Assistant Professor Greg Teegarden. This work is published: Teegarden, G.J., Cembella, A.D., Capuano*, C.L., Barron*, S.H., and Durbin, E.G. (2003) Phycotoxin accumulation in zooplankton feeding on Alexandrium fundyense - vector or sink? Journal of Plankton Research 25: 429-443 (*Student authors)
(3) Michael Butler (’02). (Independent study, spring 2002). Mike studied the biomechanics of color the feathers of ospreys under the guidance of Professor Amy Johnson and Research Associate in Mathematics and Biology Olaf Ellers. This work is published: http://www.ncbi.nlm.nih.gov/pubmed/14668312?dopt=Abstract. Mike is now in graduate school at Arizona State University.
(4) Michelle Weaver (‘03). (Gibbons Fellow, Summer 2001, 2002). Michelle worked on the Coastal Studies Center Marine Organism database under the guidance of Laboratory Instructory David Guay.
(5) Jeff Rubens (‘03). (Hughes Fellow, Summer 2001). Jeff studied the impacts of trematode parasites on growth, survivorship, and anti-predator defenses of the blue mussel Mytilus edulis under the guidance of of the first Coastal Studies Center Scholar-in-Residence Dean McCurdy
(6) Todd Forsgren (‘03). (Doherty Fellow, Summer 2001). Todd studied microparasites and sex-ratio biases in populations of intertidal amphipods under the guidance of the first Coastal Studies Center Scholar-in-Residence Dean McCurdy and Professor Mike Palopoli.
(7) Bridgid Moran (‘02). (Hughes Fellow, Summer 2001; Independent study, Fall 2001). Bridgid studied parasitism of the predatory worm Nereis virens to address the question: can parasites influence community structure? under the guidance of the first Coastal Studies Center Scholar-in-Residence Dean McCurdy and Professor Amy Johnson
(8) Michael Pesa-Fallow (‘02). (Doherty Fellow, Summer 2001). Michael studied the impacts of trematode parasites on growth, survivorship, and burying behavior of the soft-shell clam Mya arenaria under the guidance of the first Coastal Studies Center Scholar-in-Residence Dean McCurdy.
SECOND SUMMER
Summer 2000/Academic Year 2000-2001; (Johnson sabbatical thru summer 2001)
(1) Anjali Dotson (‘04). (Independent study, Academic year 2000-2001). Anjali studied zooplankton feeding behavior and selectivity based on Alexandrium spp. Toxicity under the guidance of visiting Assistant Professor Greg Teegarden
(2) Angela Dubois (’01). (Independent study, Academic year 2000-2001). Angela studied toxic algal species under the guidance of visiting Assistant Professor Greg Teegarden.
(3) Erin Giggey (’01). (Independent study, Academic year 2000-2001). Erin studied toxic algal species under the guidance of visiting Assistant Professor Greg Teegarden.
FIRST SUMMER
Summer 1999/Academic Year 1999-2000
(1) Rachel Sanders (’00). (Doherty Scholar, Summer 1999; Senior Honors research, Academic year 1999-2000). Rachel, who was the first research student to work in the marine lab, studied growth of sea cucumbers grown at two different flow speeds in the two flow tanks in the marine lab under the guidance of Professor Amy Johnson. Senior Honors Research Title: Living materials that flow as they grow.