Miocene

Miocene Taxa

Matti Aumala
Archaeopristrix was the first known "fully modern" mosark and represents the best-preserved example of post-Cretaceous mosasaur diversity. Known from a virtually complete skeleton, Archaeopristrix is surprisingly similar to small mosarks like the sakhala and shows all the major developements that characterize today's lizardwhales, including a bent, fin-supporting tail, reduced hind flippers, and nostrils placed above the eyes.

Matti Aumala Tricerolophus chooi is the first ungulaped hadrosaur known from Europe, dating back to middle Miocene. It may well be the common ancestor, or at least close relative, of many later Eurasian hadrosaurs. Unlike the modern saurolopes of Africa, Tricerolophus had only three horns, but seems to have been a good quadrapedal runner.


Luc J. 'Aspidel' Bailly

Early paleontologists excivating in Spec's eastern Eurasiawere quite surprised to discover the scattered remains of what appeared to be a group of tiny, ant-eating therizinosaurs, with elongated tubular snouts and large digging claws. These fossils were dubbed "microsegnosaurids" and were present in the literature for years before the more detailed examination of the fossils proved that these bizzare therizinosaurs were too bizarre to be real.

Recent studies of "microsegnosaurus", the first-discovered and most complete "microsegnosaurid" discovered proved that what was thought to be a single organism was, in fact, a chimera. The "ant-eating therizinosaur" was actually an amalgum of two unrelated organisms, a small therizinosaur possibly related to today's Seculasauridae and an alvarezsaur, the last known from the Eurasian continent. The three other species of "microsegnosaurid" (all of which based upon scattered elements) proved to be either therizinosaur of alvarezsaur material, lumped together with "microsegnosaurus" because of similarities with one part or another of its patch-work skeleton.

Matti Aumala
Matti Aumala Ville Sinkkonen



Matti Aumala

Ozpec's Riversleigh deposits (in northeastern Queensland) have produced the remains of a wide variety of fossil euclasaurs. The 5-million-year-old Wabulasaurus is one the largest and most striking of these specimens. This 2-tonne animal appears to have been an early offshoot of the line leading to the living hexacorns.

While generally similar to its modern-day relative, Wabulasaurus differed in the configuration of horns, its narrow beak, and the presence of large ceratopsian-like jugal processes. The postcranial skeleton is imperfectly known but appears to have been more gracile than the modern hexacorn.


Matti Aumala The Miocene Teratodon is an early rhynchoraptoran megapredator. It isn't exacly clear why this animal became extinct, but competition from the more advanced rynchoraptorans, and the fact that Teratodon seems not to have been much of a runner may have played a part in it's demise. Several Teratodon skeletons have been found, including a mummified specimen show a row of sturmtiger-like spines on this animal's back run down to the base of the tail, which has gained Teratodon the nickname "sabertooth sturmtiger".







Both on HE and Spec the continent of Australia is known to host archaic, surprising and just plain weird forms of life. But to uncover some of the most perplexing creatures, you have to dig deep, as a group of specpaleontologists discovered. While hoping to uncover the ancestor of the rhynchoraptorans, they uncovered a startling specimen that didn't seem to make sense at all: a partial skull with fragmentary postrcania that looked like it could have belonged to a strange pachycephalosaur with a crown of spikes and two prominent fangs. The animal was named Mirificranium coronatus, meaning confusing crowned skull.

Such a find was hard swallow for several reasons. First of all, there were no known pachycephalosaur remains found in Australia prior to this, and secondly the age seemed too far removed from the last known Specworld pachycephalosaurs. Mirificranium could be dated to the Ogliocene with reasonable accuracy, while the last of the pachycephalosaurs were supposed to have become extinct early Eocene.

The tale got another twist when another, more complete skull was found from early Miocene deposits. This animal, Stegocephalosaurus baiamei (meaning creator god Baiame's roofheaded reptile) had a less flamboyant skull covered by a flat bony pad. Though there were enough similarities in the skull morphology to classify the animal as a marginocephalian, it didn't seem to belong to a pachycephalosaur after all. Instead it had a lot in common with the most basal ceratopsians!

An ulna is known from HE Australia that has been tentatively assigned to a neoceratopsian (possibly Leptoceratops), which has lead to some speculation that neoceratopsians may be of Gondwanan origin. Mirificranium coronatus was however the first ceratopsian to show up in the Australian fossil record, so if there is any connection between it and the Early Cretaceous ulna, there remains a question about where the Australian ceratopians were hiding all this time. The Mirificranium postcrania also hints that the body of the animal was probably not that of the typical neoceratopsian, but rather like that of the more basal ceratopians. If a recently discovered partial postcrania from Early Miocene turns out to belong to Stegocephalosaurus, then the stegocephalosaurs were most likely at least habitual bipeds.

There is still much more we don't know about stegocephalosaurs. We can only guess that they used their bony heads in intraspecific combat as pachycephalosaurs may have done in their time, but the purpose of the fangs is unclear. Another unanswered question is their fate. While they seem so rare in the fossil record that it is too early to make assumptions about when exactly they disappeared, they did not make it to the holocene. Their extinction is indeed a great shame, as from live stegocephalosaurs we might have learned a great deal about the evolution of marginocephalians both in our world as in spec.

A fanciful restoration of Mirificranium, skull based largely on Stegocephalosaurus material.

Copyright © 2001-2002 Matti Aumala, Daniel Bensen and Brian Choo
Graphic design by Matti Aumala, 2003